Glancing over, girls cage looked totally normal.
I sat down and just observed a bit longer. You know those 'Eureka!' moments? I had one. I figured it out!
This summer we had placed 12" tiles in our cages to not only help keep the ratties cooled off but to hold down the fleece. Well, you see the boys didn't like getting their testicles cold so they were all walking on tip toes with their bodies sucked up to avoid the COLD tiles. After a giggle (or two) I removed all of the tiles except for the one under food.
I dare say they like them gone from this picture I took an hour later.